“…Nitrogen isotope values in particular can be misleading. First, because of the complex nonlinear relationship between food source and consumer, it is not possible to accurately estimate the proportion of meat versus plants in the diet, since large changes in the percentage of meat are indicated only by small increases in d 15 N values (Ambrose et al, 2003;Hedges and Reynard, 2007). Second, unlike herbivores that acquire all of their protein from plant leaves, foraging humans usually eat plants for their carbohydrate content, and therefore focus on the starch-and sugar-rich storage organs of plants, such as USOs and seeds (Lee, 1979;Marlowe, 2010).…”