Mud crab, Scylla serrate (Forskal, 1775) is a crab species that lives in a mangrove habitat and a commodity export beside blue swimming crab, Portunus pelagicus (Linnaeus, 1758). Blue swimming crab has essential economic value as meat in canned or in freeze food, while mud crab can be marketed in live condition because it is more resistant to live outside of water (Efrizal et al., 2018; Zakaria and Selasih 2018). Furthermore, people enjoy eating mud crab and other crustaceans due to their good taste and containing high nutrition (Kasry 1991; Catacutan 2002; Lubis et al., 2021). Mud crabs have economic value in several countries, such as Australia, Japan, Indonesia, Taiwan, and the Philippines. These countries make mud crab one of the production targets in their fishery activities (Watanabe et al., 1996; Cristensen et al., 2004; Ward et al., 2008). According to Cholik (1999), mud crab fisheries in Indonesia were obtained from natural capture in the coastal waters, especially in mangrove or estuary forests, and from aquaculture in brackish water. With the increasing demand for crab ARTICLE INFO