“…This fossil comparative sample includes specimens of Canis arnensis Del Campana, 1913 and C. etruscus from the Italian sites of Olivola, Upper Valdarno Basin, Coste San Giacomo, and Pantalla (Cherin et al, 2014); Canis apolloniensis Koufos and Kostopoulos (1997) from Apollonia-1 (Koufos, 2018); Canis chihliensis and Canis palmidens from Yushe Basin (Rook, 1993(Rook, , 1994; remains of C. mosbachensis Soergel, 1925from Cueva Victoria, Vallparadís (Bartolini Lucenti et al, 2017, Pirro Nord (Petrucci et al, 2013), 'Ubeidiya (Martínez-Navarro et al, 2009), Untermassfeld (Sotnikova, 2001), and Zhoukoudian (Jiangzuo et al, 2018). The relevant literature on the late Villafranchian canids was inspected (Del Campana, 1913;Crusafont Pairó, 1950;Thenius, 1954;Torre, 1967Torre, , 1974Torre, , 1979Bonifay, 1971;Kurtén, 1974;Pons-Moyà and Moyà-Solà, 1978;Pons-Moyà, 1981; Koufos and Kostopoulos, 1997;Wang et al, 1999;Sotnikova, 2001;Garrido and Arribas, 2008;Lacombat et al, 2008;Tedford et al, 2009;Petrucci et al, 2013;Cherin et al, 2014;Koufos, 2014Koufos, , 2018Bartolini Lucenti and Rook, 2016;Amri et al, 2017;Mecozzi et al, 2017;Jiangzuo et al, 2018). Extant specimens housed at the AMNH, MZUF, ICP, and MG-GNM (see abbreviation below) were also used for morphological and morphometrical comparisons.…”