As one of the key components in terrestrial water cycles, actual evapotranspiration (ET) links water, energy, and carbon cycles via plant physiological activities (Fisher et al., 2017;Katul et al., 2012). Although ET is primarily composed of daytime ET (ET D ), a growing body of observational and modeling evidence suggests that nighttime ET (ET N ) is also important for various ecohydrological and physiological processes from local to global scales (Forster, 2014;Padrón et al., 2020;Whitley et al., 2013). Globally, Padrón et al. (2020) showed that the average ET N /ET value was about 6.3% when calculated from the FLUXNET2015 data set and 7.9% from simulation results of global models. Moreover, ET N can exert significant impacts on the estimation of global water and carbon budgets, particularly in arid and semiarid regions (Lombardozzi et al., 2017;Resco De Dios et al., 2015), since the consideration of nighttime water use as indicated by ET N can noticeably reduce water use efficiency (WUE, defined as the ratio of CO 2 assimilation rate over water vapor loss) (Chaves et al., 2016). At field scales, neglecting ET N can cause the underestimation of total daily ET and the incomplete closure of energy balance, thus resulting in underestimated water requirements for crops (Skaggs & Irmak, 2011). Therefore, in addition to the role of ET D , a better understanding of nighttime water loss from terrestrial ecosystems also bears many important implications for hydrological, climatic, and agricultural studies (