“…In more open areas, where elephants respond to poaching by aggregating, such as has been described in Murchison Falls, Queen Elizabeth and Serengeti National Parks (Laws et a/., 1970;Eltringham & Malpas, 1980;Dublin and Douglas-Hamilton, 1987 and 87 per cent of the adult females have breasts (Poole, 1989b). Since female elephants start to develop breasts after about 9 months of pregnancy (Moss, 1988) and then, typically, continue to suckle one calf until the birth of the next (Lee and Moss, 1986), the absence of breasts indicates either delayed first reproduction or unusually long inter-calf intervals.…”