Abstract. Much work has been done on the study of vertebrate gaits over the past
several decades and efforts undertaken to apply this to fossil tracks,
especially dinosaurs and mammals such as cats, dogs, camels, and horses.
This work seeks to expand upon such studies and in particular to study
footprints laid down in sand by modern horses and apply such studies to
determine the gaits of fossil horse trackways. It thus builds upon the work
of Renders (1984a, b) and Kienapfel et al. (2014) and suggests additional
measurements that can be taken on horse footprints. In this study the
footprints left in the sand by 15 horses of various breeds with various
gaits were videotaped, photographed, described, and measured in order to
determine characteristics useful in distinguishing gaits. These results were
then applied to two new sets of fossil footprints, those of the middle
Miocene merychippine horse Scaphohippus intermontanus that I personally examined and measured and
those from the late Pleistocene horse Equus conversidens, previously illustrated and described
in the literature (McNeil et al., 2007). The latter horse exhibits a fast
gallop of around 9.4 m/s, but it is the former whose footprints are quite
unique. The quantitative and visual features of these prints are suggestive
of a medium-fast gait involving apparent “understepping” of diagonal
couplets and hind feet that overlap the centerline. The gait that most
closely matches the footprints of Scaphohippus is the “artificial” gait of a slow
rack or tölt, or pace, around 1.9 m/s, though an atypical trot of a horse
with major conformation issues or which is weaving (swaying) from side to
side is a less likely possibility. This intimates, along with the earlier
study of Renders (1984a, b), who found the artificial gait of the running walk
displayed by Pliocene hipparionine horses, that ancient horses possessed a
much greater variety of gaits than modern horses and that over time they
lost these abilities with the exception of certain gaited breeds.