“…Important climate changes related to an increase in the global temperature (Zachos et al, 2008), eustatic sea-level fluctuations (Miller et al, 2005), and large scale geotectonic events leading to the main uplift of the Andes (Ramos and Alemán, 2000), left a strong imprint on the southern biota Odreman Rivas, 1971, Pascual et al, 1996;Ortiz-Jaureguizar and Cladera, 2006). General information of this epoch largely comes from Argentinian Patagonia, where widespread marine and continental sedimentary deposits represent the main component of the Neogene stratigraphic succession (Feruglio, 1938(Feruglio, , 1944(Feruglio, , 1949a(Feruglio, , 1949b(Feruglio, , 1950Hatcher, 1900;Pascual et al, 1996;Nullo and Combina, 2002;Malumián and Náñez, 2011). Outcrops of contemporaneous Early Miocene rocks in southern Chubut and the Santa Cruz Province of Argentina have yielded a wealth of beautifully preserved fossils, making this epoch also a cornerstone of the many evolutionary hypotheses proposed since the pioneering work of the well-known Argentinean palaeontologists Florentino and Carlos Ameghino at the end of the 19 th century (Ameghino, 1887(Ameghino, , 1889(Ameghino, , 1900(Ameghino, -1902(Ameghino, , 1902(Ameghino, , 1904(Ameghino, , 1906.…”