“…In conifer plantations, thinning usually increases species diversity, particularly during younger stages (Thomas et al, 1999;Ito et al, 2006;Lindgren et al, 2006;Utsugi et al, 2006;Wilson and Puettmann, 2007;Ishii et al, 2008;Widenfalk and Weslien, 2009;Ares et al, 2010). Furthermore, several thinning experiments have demonstrated that species diversity usually increases monotonically with the level of thinning (Thomas et al, 1999;Parker et al, 2001;Moya et al, 2009; but see Ares et al (2010)); however, intensive thinning or clear cutting occasionally facilitates the dominance of one or a few understorey species, due to the dominance of herbs and shrubs that strongly inhibit the regeneration of hardwoods, thereby reducing understorey diversity (Alaback and Herman, 1991;Ito et al, 2006;Nagai and Yoshida, 2006;Sabo et al, 2009). Together, these studies suggest that an optimal thinning intensity exists for maximising species diversity in conifer plantations (see Jobidon et al, 2004), although the mechanism by which diversity is maximised has not yet been fully elucidated, primarily due to the lack of monitoring during the process of hardwood recruitment at different thinning intensities.…”