Bed surface particle size patchiness may play a central role in bedload and morphologic response to changes in sediment supply in gravel-bed rivers. Here we test a 1-D model (from Parker ebook) of bedload transport, surface grain size, and channel profile with two previously published flume studies that documented bed surface response, and specifically patch development, to reduced sediment supply. The model over predicts slope changes and under predicts average bed surface grain size changes because it does not account for patch dynamics. Field studies reported here using painted rocks as tracers show that fine patches and coarse patches may initiate transport at the same stage, but that much greater transport occurs in the finer patches. A theory for patch development should include grain interactions (similar size grains stopping each other, fine ones mobilizing coarse particles), effects of boundary shear stress divergence, and sorting due to cross-stream sloping bed surfaces.