In semiarid ecosystems, the transfer of water, sediments, and nutrients from bare to vegetated areas is known to be crucial to ecosystem functioning. Rainfall simulation experiments were performed on bare-soil and vegetated surfaces, on both wet and dry soils, in semiarid shrub-steppe landscapes of SE Spain to investigate the spatial and temporal factors and interactions that control the fi ne-scale variation in water infi ltration, runoff and soil loss, and hence the water and sediment fl ows in these areas. Three types of shrub-steppe landscapes varying in plant community and physiography, and four types of plant patches (oak shrub, subshrub, tussock grass, and short grass mixed with chamaephytes) were studied. Higher infi ltration and lower runoff and soil loss were measured on vegetation patches than on bare soils, for both dry and wet conditions. The oak-shrub patches produced no runoff, while the subshrub patches showed the highest runoff and soil loss. Despite these differences among patch types, the infl uence of vegetation patch type on the variables analysed was not signifi cant. The response of bare soil surfaces clearly varied between landscape types, yet the differences were only relevant under dry soil conditions. Stone cover, particularly the cover of embedded stones, and crust cover, were the key explanatory variables for the hydrological behaviour of bare soils. The study documents quantitatively how bare soils and vegetation patches function as runoff sources and runoff sinks, respectively, for a wide range of soil moisture conditions, and illustrates that landscape-type effects on bare-soil runoff sources may also exert an important control on the site hydrology, while the role of the vegetation patch type is less important. The effects of the control factors are modulated by antecedent soil moisture, with dry soils showing the most contrasting soil water infi ltration between landscapes and surface types.