“…The eight forests are located in Wuzhishan National Nature Reserve (WZS), Bawangling National Nature Reserve (BWL), Yinggeling National Nature Reserve (YGL), Diaoluoshan National Nature Reserve (DLS), Jianfengling National Nature Reserve (JFL), Limushan Nature Reserve (LMS), Baishiling Nature Reserve (BSL), and Ganshiling Nature Reserve (GSL) (Figure ). We took the proportion of natural primary forest area occupying the whole land area in each region as a measure of the degree of human disturbance, and assumed that the greater proportion of natural primary forest means the smaller degree of human disturbance, and vice versa (FAO, ; Sabatini et al, ) (Table ). In light of the human disturbance intensity, we categorized D. boisianus in these eight forests into three groups, including low disturbance intensity group (WZS), moderate disturbance intensity group (YGL, BWL, JFL, and DLS), and high disturbance intensity group (LMS, BSL, and GSL).…”