We used the aqueous extract from Artemisia campestris ssp. caudata to investigate its effects on the colonization of sand dune grass roots by mycorrhizal fungi and seedling growth. The percent colonization decreased with higher extract concentrations, and growth of three grass species was inhibited. Colonization by mycorrhizal fungi was more sensitive to the extract than was seedling growth, and no significant differences in the latter were found between the mycorrhizal and non-mycorrhizal treatments.
Beaches and associated dunes are constituted of unconsolidated materials, such as sand, and thus are low-strength land forms less robust than rocky cliffs (van der Meulen et al. 1991). It is estimated that 70% of sand-based coastlines in the world are presently subject to erosion (Bird 1985; Wind and Peerbolte 1993). However, natural dune systems are inclined to adjust after stress without permanent damage (Brown and McLachlin 2002), and when stabilized by plant cover they offer a first line of coastal defence against assault from wave action (Wind and Peerbolte 1993; Broadus 1993; De Ronde 1993). Natural self-sustaining dune systems interact with the sea and closely reflect changes in sea levels. At any given time no single sea level characterizes all oceans, that is, the resting position of the ocean surface, or geoid, is not uniformly elevated over the earth. Eustatic sea levels, free of influence from tides, waves and storms, thus vary from place to place as well as over time. Satellite altimetry, which permits more accurate as well as more numerous observations than older tide-gage methods of measuring sea levels, shows that the ocean is actually a spheroid modified by depressions and elevations. For example, in parts of the Indian Ocean sea levels are as much as 70 m lower than the global mean and in the North Atlantic 80 m higher (Carter 1988). Climate is governed by long-term periodic variations in the earth’s orbit that effect changes in solar radiation and, consequently, also in sea levels (Bartlein and Prentice 1989; Woodroffe 2002). As a result, ice ages repeatedly alternate with periods of interglacial warming in which ice masses contract and sea levels increase. Most of the time that has passed since the Cambrian period—approximately 500 million years—sea levels, although fluctuating on several timescales, have been higher than they are today. Because of the difficulties in documenting conditions so far in the distant past estimates of these sea levels vary considerably, but those shown in Fig. 13.1, based on different kinds of evidence, are representative of attempts at reconstruction (personal communication RA Rohde 2008).
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