In mid to high latitudes glacial and interglacial cycles have repeatedly changed the area available for plant growth. The speed at which plants are able to colonize areas at the onset of an interglacial is hypothesized to limit their distribution ranges even today (migrational lag). If the spread of plants would have been generally slow then plant diversity in previously glaciated areas would be expected to increase over time. We explore this hypothesis using results from six palynological investigations from two previously glaciated regions: central Sweden and north-eastern Germany. Rarefaction, slope of rank order abundance, and taxa accumulation plots were used to evaluate richness and evenness in pollen data in an attempt to separate richness from evenness. These analyses show little change in palynological richness for the northern sites throughout the Holocene. In contrast, the southern sites show an increase in richness and evenness during the early Holocene; this may be explained by the different initial conditions at the onset of the Holocene. A strong rise in palynological richness around 6000 and 1000 years ago at the southern sites can be attributed to the regional initiation of agriculture and major opening of the forest, respectively. For the northern sites there is no evidence for increased taxonomic diversity through time that could be due to delayed immigration of species.
Palynology provides the opportunity to make inferences on changes in diversity of terrestrial vegetation over long time scales. The often coarse taxonomic level achievable in pollen analysis, differences in pollen production and dispersal, and the lack of pollen source boundaries hamper the application of diversity indices to palynology. Palynological richness, the number of pollen types at a constant pollen count, is the most robust and widely used diversity indicator for pollen data. However, this index is also influenced by the abundance distribution of pollen types in sediments. In particular, where the index is calculated by rarefaction analysis, information on taxonomic richness at low abundance may be lost. Here we explore information that can be extracted from the accumulation of taxa over consecutive samples. The logtransformed taxa accumulation curve can be broken up into linear sections with different slope and intersect parameters, describing the accumulation of new taxa within the section. The breaking points may indicate changes in the species pool or in the abundance of high versus low pollen producers. Testing this concept on three pollen diagrams from different landscapes, we find that the break points in the taxa accumulation curves provide convenient zones for identifying changes in richness and evenness. The linear regressions over consecutive samples can be used to interand extrapolate to low or extremely high pollen counts, indicating evenness and richness in taxonomic composition within these zones. An evenness indicator, based on the rank-order-abundance is used to assist in the evaluation of the results and the interpretation of the fossil records. Two central European pollen diagrams show major changes in the taxa accumulation curves for the Lateglacial period and the time of human induced land-use changes, while they do not indicate strong changes in the species pool with the onset of the Holocene. In contrast, a central Swedish pollen diagram shows comparatively little change, but high richness during the early Holocene forest establishment. Evenness and palynological richness are related for most periods in the three diagrams, however, sections before forest establishment and after forest clearance show high evenness, which is not necessarily accompanied by high palynological richness, encouraging efforts to separate the two.
An interdisciplinary study was carried out to trace the hydrological changes of the Havel River in northeastern central Europe over the last ca. 2000 years. This research was driven by the hypothesis that the present‐day riverscape is to a large degree a result of medieval and modern human transformation of the drainage system. The river system forms a series of dammed lakes and river sections that were greatly altered through hydraulic engineering in the past. Along the middle course of the Havel, 16 sedimentary sequences available for geoarchaeological and paleoecological research were analyzed in order to reconstruct regional water level dynamics. Chronological control was ensured through a multitude of palynological, dendrochronological, archaeological, and radiocarbon data. The sections upriver from the Brandenburg/H. and Spandau weirs, representing sites with historic watermills, reveal substantial water level changes during the late Holocene. Generally, lower water levels before and higher levels during the medieval German colonization of that area (ca. A.D. 1180/1250) can be inferred. This water level increase, which is primarily attributed to dams constructed for watermills and secondarily due to a multitude of fish weirs, took place rapidly and amounted to a relative height of ca. 1.5 m. It enlarged the river's cross‐sections and increased the size of existing lakes or initiated secondary lakes that previously aggraded, and thus caused flooding of large parts of land. The rising water level even influenced the settlement topography to a large degree. Several medieval rural settlements were abandoned due to flooding. In total, a ca. 150‐km long dammed lake cascade was formed.
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