We examined tree diameter growth in 20 plots subjected to various disturbance intensities (natural, low, moderate and intensive logging) in a bid to understand the general tree growth responses in relation to habitat characteristics in subtropical rainforests of north-eastern New South Wales, Australia. Species-specific regeneration strategy, maximum size and level of shade tolerance were used to classify species into 5 groups; emergent and shade tolerant main canopy (group 1), shade tolerant mid canopy (2), shade tolerant understoreys (3), moderate shade tolerant (4), and shade intolerant (5) tree species. Data series for trees ≥ 10 cm diameter at 1.3 m above the ground level (dbh) providing observations spanning over 36-years were used in multilevel regression analyses. The results showed that spatial and temporal effects in tree growth at the stand-level are a combination of the differences between species functional-group compositions and environmental gradients. High growth responses were observed in the shade intolerant species while increasing level of shade tolerance and decreasing maximum size decreased trees growth rates. Tree growth increased with altitude on a large scale across regions, and with disturbance intensity on a small scale at the plot (stand) level. Increase in northness (south through flat to north facing sites) increased growth in species group 1 for trees < 67 cm dbh, but beyond this dbh threshold the opposite was true. These showed that saplings of species group 1 may require increased illumination to reach the forest canopy, but once in the canopy, low soil water availability may be limiting to tree growth in the north facing sites. Decrease in northness was associated with increased growth in species group 2 indicating that reduced illumination and improved soil moisture in the south facing sites were conducive for maximum growth in this species group. Maximum growth potential in species group 4 and 5 increased with decrease in eastness, suggesting that the increased afternoon solar radiation and temperature were conducive for high growth rates in these species. Although topographic gradient may determine the spatial and temporal variations in tree growth where growth appeared to increase from the crest down the slope into the creek, its effects on soil fertility and water availability, and interactions between these and other factors may make it difficult to discern clear growth patterns.
The effect of planting density on branch development was examined in 4-year-old Eucalyptus pilularis Sm. and Eucalyptus cloeziana F. Muell. plantations located near the coast of north-eastern NSW. Branch diameter, angle and status (live or dead) were measured along the entire stem of trees established at 1250, 1667 and 3333 stems per hectare (sph). Measurements of tree height and stem diameter at breast height over bark (DBH) were also recorded. Results showed that with an increase in initial planting density from 1250 to 1667 sph, branch size decreased, branch mortality on the lower stem increased, branch angle became more acute and DBH decreased in trees of both E. pilularis and E. cloeziana. A further increase in initial planting density from 1667 to 3333 sph did not significantly reduce branch size or branch angle but did result in increased branch mortality and decreased DBH in both species. These results suggest that increasing initial planting density from 1250 to 1667 sph will improve early branch control. However, there is no advantage in establishing trees at 3333 sph rather than 1667 sph to reduce branch size or increase branch mortality in either species. Clearwood production on the lower stem in all stocking treatments of both species was negligible at age 4. #
Pruning of live branches is performed to enhance wood quality in plantation trees. Stem, branch and wood properties may be altered following pruning from below, but the extent and duration of such effects are unknown for many plantation Eucalyptus species. The effect of 0%, 20%, 50%, and 70% green crown length removal on growth and wood quality was examined in 3.5-year-old plantation-grown Eucalyptus pilularis Sm. (blackbutt) and Eucalyptus cloeziana F. Muell. (Gympie messmate) trees on two subtropical sites in eastern Australia. Growth increment of the stem at breast height was not affected by 20% crown removal. However, stem increment at breast height was reduced for up to 8 months with 50% crown removal and up to 12 months with 70% crown removal. Pruning had no impact on height growth. However, the height of trees pruned to 70%, relative to the height of the surrounding unpruned trees, was temporarily reduced 12 months after pruning in both species. Stem form, stem taper, wood density, and residual branch growth above the pruned zone were unaffected by pruning. Owing to the fast vertical crown expansion, trees were able to rapidly compensate for leaf area removal without long-term reductions in stem growth. The implications for management operations utilizing pruning as a means of enhancing timber quality are discussed.
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