The base temperature (T b) and thermal time (Tt) requirements for germination and emergence of temperate herbage species were defined using a linear model of development rate against temperature. A T b of 4°C was found for all species. The Tt requirement for germination was lower for legume than grass species and generally lowest for small seeded species. The poor relationship (R 2 = 0.42) between Tt requirements for germination and 50% field emergence indicated that germination results could not be extrapolated to predict the rate of field emergence. The Tt for 50% field emergence was highest for the small-seeded grass species cocksfoot (220°Cd) and timothy (200°Cd). The times (days and Tt) for emergence of white clover and perennial ryegrass were similar from five autumn sowing dates but 57 days after sowing on 21 March 1996 ryegrass seedlings were 12 times heavier. The implication of these results for establishment of pasture mixtures is discussed together with the need for accurate measurement of soil temperatures to assist the prediction of field emergence.

Fine root endophytes (FRE) are arbuscule-forming fungi presently considered as a single species-Glomus tenue in the Glomeromycota (Glomeromycotina)-but probably belong within the Mucoromycotina. Thus, FRE are the only known arbuscule-forming fungi not within the arbuscular mycorrhizal fungi (AMF; Glomeromycotina) as currently understood. Phylogenetic differences between FRE and AMF could reflect ecological differences. To synthesize current ecological knowledge, we reviewed the literature on FRE and identified 108 papers that noted the presence of FRE and, in some, the colonization levels for FRE or AMF (or both). We categorized these records by geographic region, host-plant family and environment (agriculture, moderate-natural, low-temperature, high-altitude and other) and determined their influence on the percentage of root length colonized by FRE in a meta-analysis. We found that FRE are globally distributed, with many observations from Poaceae, perhaps due to grasses being widely distributed. In agricultural environments, colonization by FRE often equalled or exceeded that of AMF, particularly in Australasia. In moderate-natural and high-altitude environments, average colonization by FRE (~10%) was lower than that of AMF (~35%), whereas in low-temperature environments, colonization was similar (~20%). Several studies suggested that FRE can enhance host-plant phosphorus uptake and growth, and may be more resilient than AMF to environmental stress in some host plants. Further research is required on the functioning of FRE in relation to the environment, host plant and co-occurring AMF and, in particular, to examine whether FRE are important for plant growth in stressful environments. Targeted molecular primers are urgently needed for further research on FRE.

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Biomass and nitrogen partitioning Integrated design environment Phenological and morphological development Reusable organ and function classes a b s t r a c tThe Plant Modelling Framework (PMF) is a software framework for creating models that represent the plant components of farm system models in the agricultural production system simulator (APSIM). It is the next step in the evolution of generic crop templates for APSIM, building on software and science lessons from past versions and capitalising on new software approaches. The PMF contains a top-level Plant class that provides an interface with the APSIM model environment and controls the other classes in the plant model. Other classes include mid-level Organ, Phenology, Structure and Arbitrator classes that represent specific elements or processes of the crop and sub-classes that the mid-level classes use to represent repeated data structures. It also contains low-level Function classes which represent generic mathematical, logical, procedural or reference code and provide values to the processes carried out by mid-level classes. A plant configuration file specifies which mid-level and Function classes are to be included and how they are to be arranged and parameterised to represent a particular crop model. The PMF has an integrated design environment to allow plant models to be created visually. The aims of the PMF are to maximise code reuse and allow flexibility in the structure of models. Four examples are included to demonstrate the flexibility of application of the PMF; 1. Slurp, a simple model of the water use of a static crop, 2. Oat, an annual grain crop model with detailed growth, development and resource use processes, 3. Lucerne, perennial forage model with detailed growth, development and resource use processes, 4. Wheat, another detailed annual crop model constructed using an alternative set of organ and process classes. These examples show the PMF can be used to develop models of different complexities and allows flexibility in the approach for implementing crop physiology concepts into model set up.

Plant growth, morphology and nutritive value under shade can differ between temperate grasses. Therefore, the aim of this study was to quantify the dry matter (DM) production, sward morphology, crude protein (CP%), organic matter digestibility (OMD) and macro-nutrient concentrations (P, K, Mg, Ca and S) in a grazed cocksfoot (Dactylis glomerata L.) pasture under 10-year-old Pinus radiata D. Don forest. Four levels of light intensity were compared: full sunlight (100% photosynthetic photon flux density-PPFD), open + wooden slats (~43% PPFD), trees (~58% PPFD) and tree + slats (~24% PPFD). The mean total DM production was 8.2 t DM ha -1 yr -1 in the open and 3.8 t DM ha -1 yr -1 in the trees + slats treatment. The changes in cocksfoot leaf area index (LAI) were related to variations in morphological aspects of the sward such as canopy height and tiller population. CP% increased as PPFD declined with mean values of 18.6% in open and 22.5% in the trees + slats treatment. In contrast, the intensity of fluctuating shade had little effect on OMD with a mean value of 79 AE 3.2%. The mean annual macro-nutrient concentrations in leaves increased as the PPFD level declined mainly between the open and the trees + slats treatments. It therefore appears that heavily shaded dominant temperate pastures in silvopastoral systems limit animal production per hectare through lower DM production rates and per animal through reduced pre-grazing pasture mass of lower bulk density from the etiolated pasture.