Highlights d Genome-wide analysis of 110 ancient individuals from the Near East d Gene pools of Anatolia and Caucasus were biologically connected 6500 BCE d Gene flow from neighboring populations in Northern Levant during 3 rd millennium BCE d One individual of likely Central Asian origin in 2 nd millennium BCE Northern Levant

Archaeological and archaeogenetic evidence points to the Pontic–Caspian steppe zone between the Caucasus and the Black Sea as the crucible from which the earliest steppe pastoralist societies arose and spread, ultimately influencing populations from Europe to Inner Asia. However, little is known about their economic foundations and the factors that may have contributed to their extensive mobility. Here, we investigate dietary proteins within the dental calculus proteomes of 45 individuals spanning the Neolithic to Greco-Roman periods in the Pontic–Caspian Steppe and neighbouring South Caucasus, Oka–Volga–Don and East Urals regions. We find that sheep dairying accompanies the earliest forms of Eneolithic pastoralism in the North Caucasus. During the fourth millennium bc, Maykop and early Yamnaya populations also focused dairying exclusively on sheep while reserving cattle for traction and other purposes. We observe a breakdown in livestock specialization and an economic diversification of dairy herds coinciding with aridification during the subsequent late Yamnaya and North Caucasus Culture phases, followed by severe climate deterioration during the Catacomb and Lola periods. The need for additional pastures to support these herds may have driven the heightened mobility of the Middle and Late Bronze Age periods. Following a hiatus of more than 500 years, the North Caucasian steppe was repopulated by Early Iron Age societies with a broad mobile dairy economy, including a new focus on horse milking.

Abstract

Recently declassified photographs taken by U2 spy planes in the 1950s and 1960s provide an important new source of historical aerial imagery useful for Eurasian archaeology. Like other sources of historical imagery, U2 photos provide a window into the past, before modern agriculture and development destroyed many archaeological sites. U2 imagery is older and in many cases higher resolution than CORONA spy satellite imagery, the other major source of historical imagery for Eurasia, and thus can expand the range of archaeological sites and features that can be studied from an aerial perspective. However, there are significant barriers to finding and retrieving U2 imagery of particular locales, and archaeologists have thus not yet widely used it. In this article, we aim to reduce these barriers by describing the U2 photo dataset and how to access it. We also provide the first spatial index of U2 photos for the Middle East. A brief discussion of archaeological case studies drawn from U2 imagery illustrates its merits and limitations. These case studies include investigations of prehistoric mass-kill hunting traps in eastern Jordan, irrigation systems of the first millennium BC Neo-Assyrian Empire in northern Iraq, and twentieth-century marsh communities in southern Iraq.