Harvestmen belong to the order Opiliones, and, unlike other arachnids, they are highly polygynandrous, with both males and females mating multiply throughout the breeding season. In this chapter, we review the current information on sexual selection in the group, focusing mostly on intersexual interactions. Particularly, we provide an overview of harvestman mating systems, examine different temporal phases of male-female sexual interactions, and explore cases of sex role reversal. Several traits in harvestmen make them unique in the context of most previous studies of sexual selection. First, they have evolved an intromittent organ independently of other well-studied taxa, such as insects, spiders, and mammals. Second, the lack of long-range perception mechanisms reduces the window of opportunity for males and females to exchange information during the very short period between the first contact and intromission. In some cases, however,
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