Ecosystem fire regimes are a consequence of interactions among fuels and climate. The reconstruction of past fire regimes is of great importance for studying past climate changes and controls. The aim of this study is to compare two charcoal and pollen records from the Nothofagus forest-grass steppe ecotone (50°S) and the grass steppe (49°S). We compare fire regime responses to local vegetation changes, and inferred variation on climate conditions between 49° and 52°S during the Holocene. Cerro Frías and La Tercera charcoal records seem to be sensitive to centennial and millennial timescales of vegetation and climate variability inferred for southern Patagonia. During the Pleistocene–Holocene transition and during the middle Holocene, forest patchiness provided fuel vertical and horizontal continuity favoring fire activity. In contrast, in steppe environments high cover of shrubs favored grass patchiness, impeding fire spread. Higher pollen richness and variability through time seem to be related to higher vegetation patchiness. Past fire regime variability at the eastern side of the Andes has been closely related to westerly moisture influence between 52° and 48°S. Southern latitudes have been influenced by polar air mass intrusions to the continent during the last 5000 cal. yr BP.

Southern Patagonia intersects the core of the Southern Westerlies, providing a unique opportunity for palaeo-reconstructions and the implication of past wind variations. There is a strong link between the strength of the westerlies and precipitation, which impacts vegetation communities. The palaeoenvironmental changes that occurred since the middle Holocene to present days are reconstructed from a peat record recovered in Peninsula Avellaneda (50°15′57″S; 72°50′33″W). Interpretations are based on pollen, charcoal and sediment analyses. Closed Nothofagus forest together with low fire activity characterized the region during the period c. 5500–3500 cal. yr BP, related to an increased in precipitation through an intensification of the westerlies. After 3500 cal. yr BP, shrub and grass communities became dominant at the expense of Nothofagus forest, suggesting a climatic shift to dry conditions, as a result of weaker westerlies. This patchy forest-steppe vegetation has favoured the occurrence of fire events. Subsequently, Nothofagus forest recovered for a short period between 2000 and 400 cal. yr BP. Then, major vegetation changes took place with the re-establishment of an open landscape. The calculated moisture balance index allowed us to hypothesize about the relationship between westerlies fluctuation and El Niño Southern Oscillation (ENSO) anomalies during the mid and late Holocene.

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