Prosper, a phenomenological model of water exchange between soil, plant, and atmosphere, was used to simulate evapotranspiration and annual drainage for 2 years from a mature oak-hickory forest in the southern Appalachians. The simulation was tested by comparing drainage to measured streamflow. In a year of unusually high precipitation the simulated annual drainage was within 1.5% of measured streamflow. Simulations were also performed by using the same 2 years of meteorologic data, but vegetation parameters were changed to represent a young white pine plantation and a regrowing hardwood forest I year after clear-cutting. The model estimated that drainage for an average rainfall year was reduced 20 cm by a 16-year-old white pine plantation and increased 36 cm by clear-cutting. These results were comparable to changes of -20 and +38 cm observed in watershed experiments at Coweeta Hydrologic Laboratory. Simulated evapotranspiration during the summer was nearly identical for hardwood and pine forests, while winter and early spring water loss was greater for pine. Simulation suggests that the greater evapotranspiration by pine was due to increased interception in all seasons and increased transpiration in the dormant season. For the clear-cut area, simulated evapotranspiration was considerably less than it was for the pine or hardwood forest and thus caused simulated soil moisture contents to be greater during the summer season. of the Forest Biomass, International Union of Forest Research Organizations, College of Life Sciences and Agriculture, University of Maine, Orono, 1973. Swift, L. W., Jr., Effect of forest cover and mountain physiography on : the radiant energy balance, D.F. thesis, Duke Univ., Durham, N. C., 1972. Swift, L. W., Jr., and R. J. Luxmoore, Computational algorithm for solar radiation on mountain slopes, Memo Rep. 73-10, 20 pp., East.