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The dating of the Stone Age sequence in southern Africa has been considerably revised over the last decade, and one of the anomalies which has resulted is that the Middle Stone Age, now dated to beyond 30,000 B.P., does not immediately precede the Later Stone Agesensu stricto. The excavation and analysis of occupation horizons dating between the most recent Middle Stone Age assemblages and the Holocene is therefore of particular interest. Nelson Bay Cave, situated on the southern coast of South Africa, contains deposits which partly fill the “gap” between the Middle and Later Stone Ages, and the occupation horizons dating between about 18,000 and 5000 years ago are described in this paper. Changes in the habitat in the vicinity of the site caused by sea-level and vegetation changes coincident with the amelioration of temperatures at the end of the Pleistocene are clearly marked in the faunal remains at the site. Largely correlated with the faunal changes (which includes the introduction of marine resources to the cave at about 12,000 B.P.) are changes in the stone artifact assemblages. Three industries are recognized in the sequence: the Robberg, characterized by microbladelets produced from bladelet cores and a few small scrapers and backed tools; the Albany, characterized by large scrapers and an absence of backed tools; and the Wilton, characterized by a variety of Formal Tools including relatively large numbers of small scrapers and backed tools. These changes in artifact-manufacturing traditions are interpreted as signaling adjustments to changing environmental conditions. An explanation for these adjustments is not sought in a simple cause-and-effect relationship between the environment and the cultural response; artifact changes are seen instead as the result of a twofold process, with the environment acting as an external stimulus to change, and the direction of the artifact change governed by the selection of a range of possibilities offered by the technology of the Later Stone Agesensu latothat was widespread in subequatorial Africa during the last 20,000 years.

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