The dispersal of early humans from Africa by 1.75 Myr ago led to a marked expansion of their range, from the island of Flores in the east to the Iberian peninsula in the west. This range encompassed tropical forest, savannah and Mediterranean habitats, but has hitherto not been demonstrated beyond 45 degrees N. Until recently, early colonization in Europe was thought to be confined to the area south of the Pyrenees and Alps. However, evidence from Pakefield (Suffolk, UK) at approximately 0.7 Myr indicated that humans occupied northern European latitudes when a Mediterranean-type climate prevailed. This provided the basis for an 'ebb and flow' model, where human populations were thought to survive in southern refugia during cold stages, only expanding northwards during fully temperate climates. Here we present new evidence from Happisburgh (Norfolk, UK) demonstrating that Early Pleistocene hominins were present in northern Europe >0.78 Myr ago when they were able to survive at the southern edge of the boreal zone. This has significant implications for our understanding of early human behaviour, adaptation and survival, as well as the tempo and mode of colonization after their first dispersal out of Africa.

The colonization of Eurasia by early humans is a key event after their spread out of Africa, but the nature, timing and ecological context of the earliest human occupation of northwest Europe is uncertain and has been the subject of intense debate. The southern Caucasus was occupied about 1.8 million years (Myr) ago, whereas human remains from Atapuerca-TD6, Spain (more than 780 kyr ago) and Ceprano, Italy (about 800 kyr ago) show that early Homo had dispersed to the Mediterranean hinterland before the Brunhes-Matuyama magnetic polarity reversal (780 kyr ago). Until now, the earliest uncontested artefacts from northern Europe were much younger, suggesting that humans were unable to colonize northern latitudes until about 500 kyr ago. Here we report flint artefacts from the Cromer Forest-bed Formation at Pakefield (52 degrees N), Suffolk, UK, from an interglacial sequence yielding a diverse range of plant and animal fossils. Event and lithostratigraphy, palaeomagnetism, amino acid geochronology and biostratigraphy indicate that the artefacts date to the early part of the Brunhes Chron (about 700 kyr ago) and thus represent the earliest unequivocal evidence for human presence north of the Alps.

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Considerable debate surrounds the age of the Middle Pleistocene glacial succession in East Anglia following some recent stratigraphical reinterpretations. Resolution of the stratigraphy here is important since it not only concerns the glacial history of the region but also has a bearing on our understanding of the earliest human occupation of north-western Europe. The orthodox consensus that all the tills were emplaced during the Anglian (Marine Isotope Stage (MIS) 12) has recently been challenged by a view assigning each major till to a different glacial stage, before, during and after MIS 12. Between Trimingham and Sidestrand on the north Norfolk coast, datable organic sediments occur immediately below and above the glacial succession. The oldest glacial deposit (Happisburgh Till) directly overlies the 'Sidestrand Unio-bed', here defined as the Sidestrand Hall Member of the Cromer Forest-bed Formation. Dating of these sediments therefore has a bearing on the maximum age of the glacial sequence. This paper reviews the palaeobotany and describes the faunal assemblages recovered from the Sidestrand Unio-bed, which accumulated in a fluvial environment in a fully temperate climate with regional deciduous woodland. There are indications from the ostracods for weakly brackish conditions. Significant differences are apparent between the Sidestrand assemblages and those from West Runton, the type site of the Cromerian Stage. These differences do not result from contrasting facies or taphonomy but reflect warmer palaeotemperatures at Sidestrand and a much younger age. This conclusion is suggested by the higher proportion of thermophiles at Sidestrand and the occurrence of a water vole with unrooted molars (Arvicola) rather than its ancestor Mimomys savini with rooted molars. Amino acid racemisation data also indicate that Sidestrand is significantly younger than West Runton. These data further highlight the stratigraphical complexity of the 'Cromerian Complex' and support the conventional view that the Happisburgh Till was emplaced during the Anglian rather than the recently advanced view that it dates from MIS 16. Moreover, new evidence from the Trimingham lake bed (Sidestrand Cliff Formation) above the youngest glacial outwash sediments (Briton's Lane Formation) indicates that they also accumulated during a Middle Pleistocene interglacial -probably MIS 11. All of this evidence is consistent with a short chronology placing the glacial deposits within MIS 12, rather than invoking multiple episodes of glaciation envisaged in the 'new glacial stratigraphy' during MIS 16, 12, 10 and 6.

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