In the Wyoming Basin, archaeological sites dating from the Early Archaic to Late Prehistoric are often found associated with or adjacent to dense populations of Cymopterus bulbosus (springparsley), a nutritious geophyte that would have been an important food source for prehistoric humans living in the region. Experimental data have shown that the caloric return rates of C. bulbosus were enough to support seasonal exploitation by foragers, yet there has been no direct evidence for the use of this geophyte from the archaeological record. In this study, we examine starch granules from 10 ground stone tools excavated from two stratified, multicomponent archaeological sites in the Wyoming Basin to determine if C. bulbosus was collected and consumed in the past. Taproots of C. bulbosus were collected from two populations in the immediate vicinity of the archaeological sites in order to develop a modern starch reference. Identification of Cymopterus starch granules is based on a systematic study of those reference granules. The presence of Cymopterus starch on the ground stone artifacts suggests that prehistoric foragers were collecting and consuming these geophytes. These findings support previous hypotheses about geophyte use in southern Wyoming and therefore have implications for increasing human populations as well as settlement and subsistence decisions.
Starch-rich plants have played an important role in human evolution and societal development. Collected, grown, and consumed to support ever-increasing populations, such plants are integral to understanding past human diets. With the advent of starch granule analysis, plant resources that were invisible in the archaeological record can now be revealed in the cracks and crevices of artifacts. Widespread application of this technique, however, has stalled due to a lack of rigorous and standardized protocols. For example, taxonomic identification of starch granules using consistent diagnostic characteristics is still a challenge as there are no comprehensive surveys across important (i.e., dietary) plant taxa, especially at the levels of families, genera, and species. This study provides characteristics for identifying starch granules of seven major North American plant families (Amaranthaceae, Apiaceae, Fagaceae, Liliaceae, Pinaceae, Poaceae, and Solanaceae) based on systematic, morphometric studies of modern reference materials. A dichotomous key to starch granules of the seven families was also generated to aid in identification of those from archaeological contexts. Although we have focused on plants from western North America, these families occur across the globe and have had dietary significance throughout prehistory.
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