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Measurements of rates of growth and senescence of leaf lamina per tiller and of changes in tiller population densities were made in three experiments designed to investigate the influence of sward state on leaf turnover and net production under continuous stocking.In each experiment initially uniform swards were fenced to provide four plots on which animal numbers were adjusted twice weekly to give a series of swards maintained as nearly as possible in a steady state with respect to sward surface height (range I1-6-4 cm) and herbage mass (range 440-2690 kg

Growth measurements were made on Lolium /jerenoe-dominant vegetative swards during 4-week regrowth periods following grazing or cutting.In the grazing study, the ryegrass (Lolium perenne) swards were grazed for 12-d periods with stock numbers set according to initial herbage allowances per animal of 16, 32 and 64 kg organic matter per sheep. One area was grazed in summer and another area in the autumn.In the cutting experiment four ryegrass swards growing in boxes were prepared by cutting to either 2 or 6 cm at either 14-or 28-d intervals. The swards were cut back in autumn to 2 or 6 cm. During the regrowth period half the boxes were retained in full light and half shaded to 48% of incident light.Herbage weight, rates of leaf extension, senescence, leaf appearance and tillering, water soluble carbohydrate reserve levels in tiller bases and, in the cutting experiment, sheath tube lengths and lamina lengths of leaves which had grown through the sheath tube were measured at intervals during Ihe regrowth period.Final herbage weights reflected management history but net herbage accumulation over the four-week regrowth period, though affected by season, pretreatment and light level, was similar on swards subjected to different grazing or cutting treatments.This result was partially explained by changes in the balance between growth and senescence per tiller as herbage weight increased with time during regrowth. and partially by the reduced production per tiller in harder-grazed swards being offset by a rapid increase in tiller numbers.There was a positive curvilinear relationship between lamina length and the length of the sheath tubes through which the laminae emerged.Carbohydrate reserve levels declined rapidly after grazing or cutting, the extent of the decline being related to the severity of the defoliation. Recovery commenced after four days in summer and was complete within 3 weeks but reserve levels remained low in plants grazed or cut in autumn.The need is discussed for more detailed descriptions of sward states in management studies and for more infonnation on component processes of growth, the factors affecting them and how these interact with different kinds of management systems.

A thorough understanding of the rate of depletion of ungerminated seeds in soil is necessary to understand and model the population dynamics of many plant species. To assess how edaphic conditions influence seed survival over time a long-term field study was set up. Mesh bags of seeds of 12 species were buried under 12 contrasting semi-natural and grassland habitats and retrieved at intervals over 10 years. Seed survival and viability were assessed through germination trials and chemical staining. There were clear differences in the rate of depletion of ungerminated seed between species and also differences in the variability of this measure between habitats. Seed survival was longer in soils with a higher pH, lower moisture content and lower soil C:N. Soil characteristics need to be taken into account within studies of plant populations that depend on regeneration from seed, particularly for species where seed survival is sensitive to edaphic conditions. Ignoring this influence of the dynamics of seeds under different soil conditions may have a serious impact on the success of population modelling.

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