Eoexechia gallica gen. n. and sp. n., oldest record of the fungus gnat tribe Exechiini, is described from the Lowermost Eocene amber of France. It falls as sister group of all the extant Exechiini after its addition to the morphological phylogenetic analysis of the tribe. Its discovery suggests a Paleocene age for the Exechiini, in accordance with the current phylogenetic dating. This new discovery will help to date the whole clade in future phylogenies.

The earliest Eocene Eochauliops longicornis gen. et sp. n., first fossil Malcidae, is described from the amber of Oise (France). Together with the previous discovery of a species of Blissidae in the same amber, it suggests that the Lygaeoidea were already rather diverse during the Paleocene, even if the Mesozoic record of this superfamily remains uncertain and scarce.

The extant fly family Sciaridae currently comprises a great quantity of fossil species (Evenhuis, 1994; Röschmann & Mohrig, 1995; fossilworks Gateway to the Paleobiology Database). Many of them were originally attributed to the genus Sciara, the oldest one being Sciara burmitina Cockerell, 1917 from the mid-Cretaceous Burmese amber. Mohrig & Röschmann (1994) revised several Sciara species described from the Middle Eocene Baltic amber and transferred them to different extant sciarid genera. From France, only compression fossils have been described by Théobald (1937), one from the Late Eocene and seven from the Oligocene. Here we describe the first Sciaridae from the Lowermost Eocene Oise amber (France). Blagoderov et al. (2010) previously described a Lygistorrhinidae sciaroid from the same amber.

The Anisopodidae Knab, 1912 is an ancient family of Diptera, with several extant genera well represented in the Paleogene. It is especially the case for the genus Mycetobia Meigen, 1818, known by several Baltic amber species (Wojton et al., 2019a, b). On the contrary, the extant paleotropical genus Mesochria Enderlein, 1910 is currently known by two fossil species from the Eocene Chinese amber and the Miocene Dominican amber (Grimaldi, 1991; Szadziewski et al., 2016). Here we report the first European species of Mesochria, from the lowermost amber of Oise (France), greatly extending the past distribution of this genus.

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