No abstract

Past failures of monocultures, caused by wind-throw or insect damages, and ongoing climate change currently strongly stimulate research into mixed-species stands. So far, the focus has mainly been on combinations of species with obvious complementary functional traits. However, for any generalization, a broad overview of the mixing reactions of functionally different tree species in different mixing proportions, patterns and under different site conditions is needed, including assemblages of species with rather similar demands on resources such as light. Here, we studied the growth of Scots pine and oak in mixed versus monospecific stands on 36 triplets located along a productivity gradient across Europe, reaching from Sweden to Spain and from France to Georgia. The set-up represents a wide variation in precipitation (456–1250 mm year−1), mean annual temperature (6.7–11.5 °C) and drought index by de Martonne (21–63 mm °C−1). Stand inventories and increment cores of trees stemming from 40- to 132-year-old, fully stocked stands on 0.04–0.94-ha-sized plots provided insight into how species mixing modifies stand growth and structure compared with neighbouring monospecific stands. On average, the standing stem volume was 436 and 360 m3 ha−1 in the monocultures of Scots pine and oak, respectively, and 418 m3 ha−1 in the mixed stands. The corresponding periodical annual volume increment amounted to 10.5 and 9.1 m3 ha−1 year−1 in the monocultures and 10.5 m3 ha−1 year−1 in the mixed stands. Scots pine showed a 10% larger quadratic mean diameter (p < 0.05), a 7% larger dominant diameter (p < 0.01) and a 9% higher growth of basal area and volume in mixed stands compared with neighbouring monocultures. For Scots pine, the productivity advantages of growing in mixture increased with site index (p < 0.01) and water supply (p < 0.01), while for oak they decreased with site index (p < 0.01). In total, the superior productivity of mixed stands compared to monocultures increased with water supply (p < 0.10). Based on 7843 measured crowns, we found that in mixture both species, but especially oak, had significantly wider crowns (p < 0.001) than in monocultures. On average, we found relatively small effects of species mixing on stand growth and structure. Scots pine benefiting on rich, and oak on poor sites, allows for a mixture that is productive and most likely climate resistant all along a wide ecological gradient. We discuss the potential of this mixture in view of climate change.

No abstract

A higher frequency of increasingly severe droughts highlights the need for short-term measures to adapt existing forests to climate change. The maintenance of reduced stand densities has been proposed as a promising silvicultural tool for mitigating drought stress. However, the relationship between stand density and tree drought susceptibility remains poorly understood, especially across ecological gradients. Here, we analysed the effect of reduced stand density on tree growth and growth sensitivity, as well as on short-term drought responses (resistance, recovery, and resilience) of Scots pine (Pinus sylvestris L.), sessile oak (Quercus petraea (Matt.) Liebl.), and ponderosa pine (Pinus ponderosa Douglas ex C. Lawson). Tree ring series from 409 trees, growing in stands of varying stand density, were analysed at sites with different water availability. For all species, mean tree growth was significantly higher under low compared with maximum stand density. Mean tree growth sensitivity of Scots pine was significantly higher under low compared with moderate and maximum stand density, while growth sensitivity of ponderosa pine peaked under maximum stand density. Recovery and resilience of Scots pine, as well as recovery of sessile oak and ponderosa pine, decreased with increasing stand density. In contrast, resistance and resilience of ponderosa pine significantly increased with increasing stand density. Higher site water availability was associated with significantly reduced drought response indices of Scots pine and sessile oak in general, except for resistance of oak. In ponderosa pine, higher site water availability significantly lessened recovery. Higher site water availability significantly moderated the positive effect of reduced stand density on drought responses. Stand age had a significantly positive effect on the resistance of Scots pine and a negative effect on recovery of sessile oak. We discuss potential causes for the observed response patterns, derive implications for adaptive forest management, and make recommendations for further research in this field.

In the last decades, studying effects of mixing tree species is increasingly important. In particular, under changing growing conditions and social requirements, investigations on mixed forest compared to mono-specific stands are of special interest, for example, stability, resilience or ecosystem services. Permanent forest research plots are a unique data source, providing the required information but being time-consuming and costly to establish. Moreover, large data sets of such plots are missing but needed for generalising any findings. Temporary research plots provide ad hoc information of its status quo and require less effort than permanent plots. Usually, such plots provide no information of the recent tree and stand characteristics. We demonstrate a new method developed under the scope of COST action FP 1206 EuMIXFOR (European Network on Mixed Forests) to estimate retrospective tree dynamics and stand characteristics. The results of validation reveal its usefulness for reconstructing 5-10 years. Thus, the method provides new potential in establishing larger networks across several countries, in particular, for studying underlying processes when comparing mono-specific with mixed forest stands.