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INTRODUCTION REVIEW OF LITERATURE 4 Historical 4 Evidence for the Compounds Moved and the Tissues Involved Using Conventional Techniques Use of P-^2 as a Tracer in Translocation Studies 13 Use of C^4 as a Tracer in Translocation Translocation of Temperature Effects on Translocation Studies on the Structure and Properties of the Phloem METHODS AND MATERIALS Studies with P 52 General methods Methods for specific experiments 46 Studies with 53 General methods Methods for specific experiments Studies with C^4 63 General methods 63 Methods for specific experiments RESULTS 66 Studies with P^2 and C^4 66 iii Page Normal distribution 66 Distribution studies using autoradiography 76 The effect of steaming on translocation 90 The effect of cold stem temperature on translocation 93 The effect of phosphorus level in treated plant and in treatment solution 99 light effects on translocation 105 The effect of dinitrophenol on translocation 106 The effect of growing tip removal on translocation 109 Studies with Ill Translocation of N'° in poplars 111

INTRODUCTION REVIEW OF LITERATURE 4 Historical 4 Evidence for the Compounds Moved and the Tissues Involved Using Conventional Techniques Use of P-^2 as a Tracer in Translocation Studies 13 Use of C^4 as a Tracer in Translocation Translocation of Temperature Effects on Translocation Studies on the Structure and Properties of the Phloem METHODS AND MATERIALS Studies with P 52 General methods Methods for specific experiments 46 Studies with 53 General methods Methods for specific experiments Studies with C^4 63 General methods 63 Methods for specific experiments RESULTS 66 Studies with P^2 and C^4 66 iii Page Normal distribution 66 Distribution studies using autoradiography 76 The effect of steaming on translocation 90 The effect of cold stem temperature on translocation 93 The effect of phosphorus level in treated plant and in treatment solution 99 light effects on translocation 105 The effect of dinitrophenol on translocation 106 The effect of growing tip removal on translocation 109 Studies with Ill Translocation of N'° in poplars 111