Supercritical pressure fluids are widely used in heat transfer and energy systems. The benefit of high heat transfer performance and the successful avoidance of phase change from the use of supercritical pressure fluids are well-known, but the complex behaviours of such fluids owing to dramatic thermal property variations pose strong challenges to the design of heat transfer applications. In this paper, the turbulent flow and heat transfer of supercritical pressure $\textrm {CO}_2$ in a small vertical tube influenced by coupled effects of buoyancy and thermal acceleration are numerically investigated using direct numerical simulation. Both upward and downward flows with an inlet Reynolds number of 3540 and pressure of 7.75 MPa have been simulated and the results are compared with corresponding experimental data. The flow and heat transfer results reveal that under buoyancy and thermal acceleration, the turbulent flow and heat transfer exhibit four developing periods in which buoyancy and thermal acceleration alternately dominate. The results suggest a way to distinguish the dominant factor of heat transfer in different periods and a criterion for heat transfer degradation under the complex coupling of buoyancy and thermal acceleration. An analysis of the orthogonal decomposition and the generative mechanism of turbulent structures indicates that the flow acceleration induces a stretch-to-disrupt mechanism of coherent turbulent structures. The significant flow acceleration can destroy the three-dimensional flow structure and stretch the vortices resulting in dissipation.

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