The effects of individual versus group learning (in triads) on efficiency of retention and transfer test performance in the domain of biology (heredity) among 70 high-school students were investigated. Applying cognitive load theory, the limitations of the working memory capacity at the individual level were considered an important reason to assign complex learning tasks to groups rather than to individuals. It was hypothesized that groups will have more processing capacity available for relating the information elements to each other and by doing so for constructing higher quality cognitive schemata than individuals if the high cognitive load imposed by complex learning tasks could be shared among group members. In contrast, it was expected that individuals who learn from carrying out the same complex tasks would need all available processing capacity for remembering the interrelated information elements, and, consequently, would not be able to allocate resources to working with them. This interaction hypothesis was confirmed by the data on efficiency of retention and transfer test performance; there was a favorable relationship between mental effort and retention test performance for the individual learners as opposed to a favorable relationship between transfer test performance and mental effort for the students who learned in groups. Collaborative learning models are based on the premise that certain types of learning are best achieved interactively rather than through a one-way transmission process (Johnston, James, Lye, & McDonald, 2000;Littleton & Häkkinen, 1999;Slavin, 1983Slavin, , 1995Veerman & Veldhuis-Diermanse, 2001;Weigel, 2002). Although collaborative learning is emerging as a promising educational approach, research on its effects on learning has been highly inconclusive (Kester & Paas, 2005). We believe that these inconclusive results have, among other things, been caused by a lack of attention to the structures constituting human cognitive architecture (Sweller, 1988;Sweller, Van Merriënboer, & Paas, 1998) when designing collaborative learning environments.Research stressing the potential of collaborative learning shows that collaborative learning environments can stimulate and/or enable learners to engage in activities that are valuable for learning. Activities such as self-directed learning, negotiating meaning (Beers, Boshuizen, & Kirschner, 2007;Kirschner, Beers, Boshuizen, & Gijselaers, 2008), verbalizing explanations, justifications and reflections, giving mutual support (Van Boxtel, Van der Linden, & Kanselaar, 2000), and developing arguments about complex problems or propositions (Munneke, Andriessen, Kanselaar, & Kirschner, 2007) have been found to facilitate the learning process. Collaborative learning has also been shown to help learners retain the learned information longer (Morgan, Whorton, & Gunsalus, 2000) and to foster their higher-order skills more than in more traditional lecture-based learning environments (Sloffer, Dueber, & Duffy, 1999). It is important to note that...