“…The Permian-Triassic mass extinction (PTME) was the most devastating biotic crisis in Earth history (Raup, 1979;Erwin, 1993;Alroy et al, 2008), both on land (Sahney and Benton, 2008) and in the oceans (Stanley, 2016;Fan et al, 2020). The extinction was driven by the eruption of the Siberian Traps (e.g., Renne et al, 1995;Wignall, 2001;Rampino et al, 2017;Green et al, 2022), which input tremendous amounts of CO 2 into the atmosphere (e.g., Kidder and Worsley, 2010;Black et al, 2015;Sobolev et al, 2015;Burgess et al, 2017;Joachimski et al, 2022;Tian and Buck, 2022), leading to a runaway greenhouse effect that made life in shallow tropical seas untenable for many organisms (Sun et al, 2012;Romano et al, 2013;Song et al, 2014). Global hothouse conditions led to sluggish oceanic circulation (Kidder and Worsley, 2004;Kiehl and Shields, 2005;Kiehl and Shields, 2010) and the depletion of oxygen in the deep ocean (Isozaki, 1997;Grasby et al, 2013;Grasby et al, 2021;Takahashi et al, 2021), as well as the impingement of anoxic water masses onto the continental shelves (e.g., Wignall and Hallam, 1992;Thomas et al, 2004;Algeo et al, 2007;Galfetti et al, 2008;Grasby and Beauchamp, 2009;Liao et al, 2010;Metcalfe et al, 2013;Pietsch et al, 2014;Lau et al, 2016;Xiao et al, 2018).…”