“…3). We expected that undershrub soil surfaces would be rougher than interspaces based on abundant literature that describes a wider range of particle and aggregate sizes and densities for undershrubs relative to interspaces (Blackburn, 1975;Roundy et al, 1978;Wood et al, 1978Wood et al, , 1982Doescher et al, 1984;Johnson and Gordon, 1988;Blackburn et al, 1990Blackburn et al, , 1992Bolton et al, 1990;Goff et al, 1993;Pierson et al, 1994Pierson et al, , 2001Pierson et al, , 2002Davies et al, 2007Davies et al, , 2009Aanderud et al, 2008;Hooker et al, 2008) as well as the greater proportion of organic matter on the undershrub soil surfaces that we characterized. Furthermore, interspaces often have a physical crust with a relatively flat, smooth appearance that is in sharp contrast, visually, to raised undershrub mounds (Blackburn, 1975;Roundy et al, 1978;Wood et al, 1978Wood et al, , 1982Doescher et al, 1984;Johnson and Gordon, 1988;Blackburn et al, 1990Blackburn et al, , 1992Bolton et al, 1990;Goff et al, 1993;Pierson et al, 1994Pierson et al, , 2001Pierson et al, , 2002Davies et al, 2007Davies et al, , 2009Aanderud et al, 2008;Hooker et al, 2008).…”