“…These include ocean circulation and stratification changes (Siegenthaler and Wenk, 1984;Watson and Garabato, 2006;Bouttes et al, 2010;Völker and Köhler, 2013), continental weathering (Munhoven, 2002), volcanism (Huybers and Langmuir, 2009;Roth and Joos, 2012), changes in the ocean's biological pump, e.g. by changes in its strength as modulated by the availability of micro-and macro-nutrients Tschumi et al, 2011;Oka et al, 2011), or changes in the export rain ratio Tschumi et al, 2011).…”