“…The Cenozoic Era (66 Ma to the present day) saw several dramatic changes of the marine and continental ecosystems (e.g., evolution of large plankton feeders such as baleen whales, shift towards cold-water, high nutrient plankton assemblages at high latitude, expansion of terrestrial mammals) major tectonic events (e.g., opening of Southern Hemisphere Oceanic gateways, shift to the 4-layer structure of the modern ocean, collision of the African-Arabian-Eurasian plates, uplift of the Alpine and Himalayan mountain belt) and global climate forcing (e.g., change from greenhouse to icehouse conditions) (Cerling, 1997;Houben et al, 2013;Norris et al, 2013;Cermeño et al, 2015;Mutz et al, 2018;Komar and Zeebe, 2021). The acceleration of Cenozoic climate cooling started after the Early Eocene Climatic Optimum (EECO; ~52-50 Ma), with temperatures ~10-12 °C warmer than the modern deep ocean, followed by the appearance and expansion of the Antarctic ice-sheets after the Eocene-Oligocene Transition (EOT; ~34 Ma) and ultimately culminating in the extensive Northern Hemisphere glaciation of the Pleistocene (~2.6-0.01 Ma; Zachos et al, 2001;Lear et al, 2008;Mudelsee et al, 2014;Abdullayev et al, 2021). This long-term transition in Earth`s climate is well documented in marine sedimentary archives, but its impact on the evolution of continental ecosystems remains poorly constrained, mainly because continuous, well preserved terrestrial records are scarce and the responses to climate change in these settings are highly complex, depending on latitude, proximity to coast and mountain ranges, position relative to climatic winds, vegetation etc.…”