The representation of visual information about letters is proposed to be highly systematic, involving not only abstract information that is invariant across type faces (or fonts), but also a number of parameters whose values are determined by the current font The system exploits regularities that are characteristic of letters and fonts by becoming tuned to the details of the font This should result in efficient letter perception when the stimuli are regular (when all of the letters are of a consistent font), but not when the stimuli are irregular (when the letters are from a variety of fonts) The prediction of faster processing with a regular font, as compared with a mixed font, was examined in three experiments requiring the recognition of four-letter strings Experiment 1 confirmed the prediction, and Experiment 2 replicated the effect with the number of "features" equated across conditions. Experiment 3 showed that the disadvantage for a mixture of fonts is related to how much the representational system must be adjusted to handle the different fonts A central issue in cognitive and perceptual psychology is how familiar objects are represented and perceived. This article concerns the representation and perception of letters, which are representative of more complex objects in that they vary considerably in appearance from instance to instance: The actual form of a letter depends on the type face, or font. Therefore, it is necessary for models of letter perception to specify how the perceptual system maps letters of different fonts onto the appropriate abstract letter codes. The purpose of the present research was twofold, first, to begin developing a new kind of model of letter perception, one that uses the idea of a structural network, and, second, to test this model against a class of simpler models by examining how perceptual representations might become systematically tuned for a particular font.A useful general approach to the recognition problem is to assume that letters are represented and perceived as sets of fea-