Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species

Abstract

Abstract. Two new species of Copris Geoffroy, 1762 are described and illustrated: Copris (subgenus incertae sedis) caobangensis sp. n. from Caobang Province (northern Vietnam) and Copris (Copris) sonensis sp. n. from Thanhhoa Province (central Vietnam). Copris (Copris) szechouanicus Balthasar, 1958 is recorded in Vietnam for the fi rst time and data on the morphology, distribution and ecology of this species are given. An updated species list and an identifi cation key for the Copris species so far known from Vietnam are presented along with detailed photographs of the poorly known species.

A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History

“…The specimens were examined with a Kyowa Optical Microscope (Model SDZ-P) and species were identified based on the following available published literature: Arrow (1931), Balthasar (1963aBalthasar ( , 1963b, Löbl & Smetana (2006), Montreuil (2011), Abbas et al (2015Abbas et al ( , 2023, Génier & Moretto (2017), Bui et al (2018), Edmonds (2018), andMinetti (2022). After examining 605 specimens of dung beetles, the digital photographs along with the following data are given for all available species housed at PMNH: country name, locality, number of examined specimens, and complete label data for 26 species in 12 genera and seven tribes: Coprini (nine species ≈35%), Onthophagini (six species ≈23%), Onitini (six species ≈23%), Gymnopleurini (two species ≈8%), Oniticellini, Scarabaeini, and Sisyphini each represented by a single species.…”