More than 500 species of the Poaceae are found in Kenya, East Africa. Eighteen of twenty-seven tribes are exclusively (except the Paniceae and Danthonieae) of the C photosynthetic type. A floristic analysis of low altitude grasslands suggests that nearly all species at these low altitudes are of the C photosynthetic type. At high altitudes, however, nearly all grasses are of the C photosynthetic type. Open grassland vegetation was sampled along a transect from arid low altitude sites to the top of Mt. Kenya in an attempt to document the general distributions of the photosynthetic types.The major tribes illustrated three general patterns of distribution. The C tribes Chlorideae, Eragrosteae, Sporoboleae, and Aristideae were abundant at low altitudes (or low indices of available soil moisture). The Paniceae and Andropogoneae were also exclusively C but were more common at intermediate altitudes. The C tribes Aveneae, Festuceae, and Agrostideae were found only at high altitudes. In these open grasslands there were no C species below 2,000 m and no C species above 3,000 m. The variation in δC of the live grass vegetation with altitude confirms these distributional patterns and suggests a sharp transition zone between these two photosynthetic types. The photosynthetic type accounts for broad distributions within the Poaceae but these distributions are further modified by characteristics which may be inherent in the tribal groups. Ecological and paleoecological significance of these patterns of distribution are discussed.
The quantitative plant species composition of the rumen contents of a large number of individuals from eight East African herbivores was determined by direct visual analysis. All plant species were classified as either C or C, and an estimated δC for the rumen sample was calculated. This estimated value was compared to a measured value determined directly from rumen subsample. The two methods of determining quantitative C and C composition differed by less than 1%, and the isotopic analysis has the advantage of being rapid and totally objective.The isotopic analysis allowed us to differentiate between grazers and browsers and to determine the quantitative dependence of each animal on C and C photosynthetic types. Kongoni, wildebeest, cattle, and sheep were nearly pure grazers on the Athi Kapiti Plains; and the Grant's gazelle were predominantly browsers. Thompson's gazelle, goast and impala were intermediate. The species most dependent upon browse showed a marked and rapid shift to grass within a few days following rain. This isotopic method may have general utility in the study of East African ecology.
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