Abstract.Little is known about the terrestrial response of high-latitude Scandinavian vegetation to the warmerthan-present climate of the late Pliocene (Piacenzian, 3.60-2.58 Ma). In order to assess Piacenzian terrestrial climate variability, we present the first high-resolution reconstruction of vegetation and climate change in northern Norway between 3.6 and 3.14 Ma. The reconstructions are derived from pollen assemblages in the marine sediments of ODP Hole 642B, Norwegian Sea (67 • N). The palynological assemblages provide a unique record of latitudinal and altitudinal shifting of the forest boundaries, with vegetation alternating between cool temperate forest during warmerthan-present intervals and boreal forest similar to today during cooler intervals. The northern boundary of the nemoral to boreonemoral forest zone was displaced at least 4-8 • further north, and warmest-month temperatures were 6-14.5 • C higher than at present during warm phases. Warm climatic conditions persisted during the earliest Piacenzian (ca. 3.6-3.47 Ma) with diverse cool temperate nemoral to boreonemoral forests growing in the lowlands of the Scandinavian mountains. A distinct cooling event at ca. 3.47 Ma resulted in a southward shift of vegetation zones, leading to the predominance of boreal forest and the development of open, low alpine environments. The cooling culminated around 3.3 Ma, coinciding with Marine Isotope Stage (MIS) M2. Warmer climate conditions returned after ca. 3.29 Ma, with higher climate variability indicated by the repeated expansion of forests and peatlands during warmer and cooler periods, respectively. Climate progressively cooled after 3.18 Ma, resembling climatic conditions during MIS M2. A high variability of Norwegian vegetation and climate changes during the Piacenzian is superimposed on a long-term cooling trend. This cooling was accompanied by an expansion of Sphagnum peatlands that potentially contributed to the decline in atmospheric CO 2 concentrations at the end of the Piacenzian warm period and facilitated ice growth through positive vegetation-snow albedo feedbacks. Correlations with other Northern Hemisphere vegetation records suggest hemisphere-wide effects of climate cooling.

Abstract. Little is known about the terrestrial response of high latitude Scandinavian vegetation to the warmer-than-present climate of the Late Pliocene (Piacenzian, 3.60–2.58 Ma). In order to assess Piacenzian terrestrial climate variability we present the first high resolution reconstruction of vegetation and climate change in northern Norway between 3.6–3.14 Ma. The reconstructions are derived from pollen assemblages in the marine sediments of ODP Hole 642B, Norwegian Sea (67° N). The palynological assemblages provide a unique record of latitudinal and altitudinal shifting of the forest boundaries, with vegetation alternating between cool temperate forest during warmer-than-present intervals, and boreal forest similar to today during cooler intervals. The northern boundary of the nemoral to boreonemoral forest zone was displaced at least 4–8° further north and warmest month temperatures were 6–14.5 °C higher than present during warm phases. Warm climatic conditions persisted during the earliest Piacenzian (ca. 3.6–3.47 Ma) with diverse cool temperate nemoral to boreonemoral forests growing in the lowlands of the Scandinavian mountains. A distinct cooling event at ca. 3.47 Ma resulted in a southward shift of vegetation boundaries, leading to the predominance of boreal forest and the development of open, low alpine environments. The cooling culminated around 3.3 Ma, coinciding with Marine Oxygen Isotope Stage (MIS) M2. Warmer climate conditions returned after ca. 3.29 Ma with higher climate variability indicated by the repeated expansion of forests and peat lands during warmer and cooler periods, respectively. Climate progressively cooled after 3.18 Ma, resembling climatic conditions during MIS M2. A high variability of Norwegian vegetation and climate changes during the Piacenzian is superimposed on a long-term cooling trend. This cooling was accompanied by an expansion of Sphagnum peat lands that potentially contributed to the decline in atmospheric CO2-concentrations at the end of the Piacenzian warm period and facilitated ice growth through positive vegetation-snow albedo feedbacks. Correlations with other Northern Hemisphere vegetation records suggest hemisphere-wide effects of climate cooling.

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During the Pliocene Epoch, a stronger-than-present overturning circulation has been invoked to explain the enhanced warming in the Nordic Seas region in comparison to low to midlatitude regions. While marine records are indicative of changes in the northward heat transport via the North Atlantic Current (NAC) during the Pliocene, the long-term terrestrial climate evolution and its driving mechanisms are poorly understood. We present the first two-million-year-long Pliocene pollen record for the Nordic Seas region from Ocean Drilling Program (ODP) Hole 642B, reflecting vegetation and climate in Arctic Norway, to assess the 2 influence of oceanographic and atmospheric controls on Pliocene climate evolution. The vegetation record reveals a long-term cooling trend in northern Norway, which might be 24 linked to a general decline in atmospheric CO2 concentrations over the studied interval, and 25 climate oscillations primarily controlled by precession (23 kyr), obliquity (54 kyr) and eccentricity (100 kyr) forcing. In addition, the record identifies four major shifts in Pliocene vegetation and climate mainly controlled by changes in northward heat transport via the NAC. Cool temperate (warmer than present) conditions prevailed between 5.03-4.30 Ma, 3.90-3.47 Ma and 3.29-3.16 Ma and boreal (similar to present) conditions predominated between 4.30-3.90 Ma, 3.47-3.29 and after 3.16 Ma. A distinct decline in sediment and pollen accumulation rates at c. 4.65 Ma is probably linked to changes in ocean currents, marine productivity and atmospheric circulation. Climate model simulations suggest that changes in the strength of the Atlantic Meridional Overturning Circulation during the Early Pliocene could have affected atmospheric circulation in the Nordic Seas region, which would have affected the direction of pollen transport from Scandinavia to ODP Hole 642B.